G.G.E. Scudder. 1998. Heteroptera in Smith, I.M., and G.G.E. Scudder, eds. Assessment of species diversity in the Montane Cordillera Ecozone. Burlington: Ecological Monitoring and Assessment Network, 1998.

HETEROPTERA

G.G.E. Scudder

APPENDIX

Systematic list of the of the Montane Cordillera Heteroptera.

(*=Potentially rare and endangered; **=Endemic; A =Alien and introduced).

Infraorder DIPSOCOMORPHA

Superfamily DIPSOCOIDEA

Family CERATOCOMBIDAE

Ceratocombus vagans McAtee & Malloch

Infraorder NEPOMORPHA

Superfamily NEPOIDEA

Family BELOSTOMATIDAE

Belostoma flumineum Say

Lethocerus americanus (Leidy)

Family NEPIDAE

Ranatra fusca Palisot

Superfamily CORIXOIDEA

Family CORIXIDAE

Arctocorisa convexa (Fieber)

A. sutilis (Uhler)

Callicorixa audeni Hungerford

C. scudderi Jansson

Cenocorixa bifida hungerfordi Lansbury

C. expleta (Uhler)

C. utahensis (Hungerford)

Corisella decolor (Uhler)

Hesperocorixa atopodonta (Hungerford)

H. laevigata (Uhler)

H. michiganensis (Hungerford)

H. vulgaris (Hungerford)

Sigara bicoloripennis (Walley)

S. conocephala (Hungerford)

S. decoratella (Hungerford)

S. penniensis (Hungerford)

S. grossolineata Hungerford

S. mathesoni Hungerford

S. mullattensis (Hungerford)

S. omani (Hungerford)

S. solensis (Hungerford)

S. washingtonensis Hungerford

Dasycorixa hybrida (Hungerford)

D. johanseni (Walley)

D. rawsoni Hungerford

Cymatia americana Hussey

Superfamily NOTONECTOIDEA

Family NOTONECTIDAE

Buenoa confusa Truxal

B. macrotibialis Hungerford

Notonecta borealis Hussey

N. kirbyi Hungerford

* N. spinosa Hungerford

N. undulata Say

* N. unifasciata andersoni Hungerford

Infraorder GERROMORPHA

Superfamily MESOVELIOIDEA

Family MESOVELIIDAE

Mesovelia mulsanti White

Superfamily HEBROIDEA

Family HEBRIDAE

Merragata hebroides White

Superfamily HYDROMETROIDEA

Family HYDROMETRIDAE

* Hydrometra martini Kirkaldy

Superfamily GERROIDEA

Family VELIIDAE

Microvelia buenoi Drake

M. pulchella Westwood

Family GERRIDAE

Aquarius nyctalis (Drake & Hottes)

A. remigis (Say)

Gerris buenoi Kirkaldy

G. comatus Drake & Hottes

G. incognitus Drake & Hottes

G. incurvatus Drake & Hottes

G. pingreensis Drake & Hottes

Limnoporus dissortis (Drake & Hottes)

L. notabilis (Drake & Hottes)

Infraorder CIMICOMORPHA

Superfamily CIMICOIDEA

Family CIMICIDAE

Cimex lectularius Linnaeus

C. pilosellus (Horvath)

Oeciacus vicarius Horvath

* Hesperocimex coloradensis List

Family ANTHOCORIDAE

Acompocoris lepidus (Van Duzee)

Anthocoris antevolens White

A. musculus (Say)

A A. nemoralis (Fabricius)

A. tomentosus Pericart

A. whitei Reuter

Elatophilus pullus Kelton & Anderson

Melanocoris longirostris Kelton

M. nigricornis Van Duzee

Tetraphleps canadensis Provancher

T. feratis (Drake & Hottes)

T. latipennis Van Duzee

T. pilosipes Kelton & Anderson

T. uniformis Parshley

Macrotracheliella nigra Parshley

Orius diespeter Herring

O. tristicolor (White)

Cardiastethus borealis Kelton

Dufouriellus ater (Dufour)

Scoloposcelis flavicornis Reuter

Xylocoris cursitans (Fallén)

X. galactinus (Fieber)

X. umbrinus Van Duzee

Family LYCTOCORIDAE

L. stalii (Reuter)

L. tuberosus Kelton & Anderson

Superfamily REDUVIOIDEA

Family REDUVIIDAE

Barce fraterna (Say)

Empicoris errabundus (Say)

E. orthoneuron McAtee & Malloch

Fitchia spinosula Stål

Pselliopus spinicollis (Champion)

Reduvius personatus (Linnaeus)

Rhynocoris ventralis (Say)

Sinea diadema (Fabricius)

Zelus longipes (Fabricius)

Z. tetracanthus Stål

Family PHYMATIDAE

Phymata americana metcalfi Evans

Family MIRIDAE

Subfamily BRYOCORINAE

Tribe DICYPHINI

Dicyphus discrepans Knight

D. hesperus Knight

D. vestitus Uhler

Macrolophus rivalis (Knight)

Tupiocoris confusus (Kelton)

T. elongatus (Van Duzee)

T. rubi (Knight)

T. similis (Kelton)

T. tibialis (Kelton)

Usingerella bakeri (Knight)

Subfamily DERAEOCORINAE

Tribe CLIVENEMATINI

Clinema medialis Knight

Largidea grossa Van Duzee

L. shoshonea Knight

Tribe DERAEOCORINI

Deraeocapsus fraternus (Van Duzee)

Deraeocoris albigulus Knight

* D. bakeri Knight

D. barberi Knight

D. brevis (Uhler)

D. diveni Knight

D. fasciolus Knight

D. fulvescens (Reuter)

* D. incertus Knight

D. kennicotti Knight

D. mutatus Knight

D. piceicola Knight

D. rubroclarus Knight

D. schwartzii (Uhler)

D. triannulipes Knight

Eurychilopterella pacifica Stonedahl

Eustictus necopinus Knight

Tribe HYALIODINI

Hyaliodes harti Knight

Subfamily MIRINAE

Tribe MIRINI

A Adelphocoris lineolatus (Goeze)

A. rapidus (Say)

A. superbus (Uhler)

Agnocoris rubicundus (Fallén)

A. utahensis Moore

A Capsus ater (Linnaeus)

C. cinctus (Kolenati)

Dichrooscytus alpinus Kelton

D. convexifrons Knight

**D. flagellatus Kelton

D. flavescens Knight

D. irroratus Van Duzee

D. latifrons Knight

D. raineri Knight

D. rostratus Kelton

D. ruberellus Knight

D. rubidus Kelton

**D. rufusculus Kelton

D. suspectus Reuter

Henrilygus nubilus (Van Duzee)

H. ultranubilus (Knight)

Irbisia brachycera (Uhler)

I. elongata Knight

I. fuscipubescens Knight

I. nigripes Knight

* I. pacificus (Uhler)

I. serrata Bliven

* I. shulli Knight

Lygidea annexa (Uhler)

L. obscura Reuter

L. rosacea Reuter

L. salicis Knight

Lygocoris alni (Knight)

L. atritylus (Knight)

L. communis (Knight)

L. contaminatus (Fallén)

L. pabulinus (Linnaeus)

Lygus atriflavus Knight

L. atritibialis Knight

L. borealis (Kelton)

L. ceanothi Knight

L. convexicollis Reuter

L. elisus Van Duzee

L. hesperus Knight

L. humeralis Knight

L. keltoni Schwartz

L. lineolaris (Palisot)

L. lupini Schwartz

L. potentillae Kelton

L. punctatus (Zetterstedt)

L. rugulipennis Poppius

L. robustus (Uhler)

L. rubroclarus Knight

L. rubrosignatus Knight

L. rufidorsus (Kelton)

L. scudderi Schwartz

L. shulli Knight

L. solidaginis (Kelton)

L. striatus Knight

L. unctuosus (Kelton)

Nonlygus nubilatus (Knight)

Neoborella xanthenes Henry

Neurocolpus longirostris Knight

N. nubilis (Say)

Orthops scutellatus Uhler

Phytocoris alpinus Kelton

P. californicus Knight

P. conspersipes Reuter

P. conspurcatus Knight

P. driesbachi Knight

P. dumicola Stonedahl

P. fraterculus Van Duzee

P. inops Uhler

P. interspersus Uhler

P. juliae Stonedahl

P. juncundus Van Duzee

P. knowltoni Knight

P. lasiomeris Reuter

P. neglectus Knight

P. pallidicornis Reuter

P. plenus Van Duzee

P. politus (Reuter)

P. purshiae Stonedahl

P. rostratus Knight

P. stellatus Van Duzee

P. strigosus Knight

P. tenuis Van Duzee

P. yollabollae Bliven

Pinalitus approximatus (Stål)

P. rostratus Kelton

P. solivagus (Van Duzee)

Platylygus luridus (Reuter)

P. piceicola Kelton

P. pseudotsugae Kelton

P. rolfsi Knight

P. rubripes Knight

Plesiocoris rugicollis (Fallén)

Polymerus basivittis pallidulus Knight

P. brevirostris Knight

P. chrysopsis Knight

P. cognatus (Fieber)

P. diffusus (Uhler)

P. rufipes Knight

P. tumidifrons Knight

P. unifasciatus (Fabricius)

P. venaticus (Uhler)

Salignus tahoensis (Knight)

A Stenotus binotatus (Fabricius)

Tribe RESTHENINI

Prepops bivittis (Stål)

P. borealis (Knight)

P. eremicola (Knight)

P. nigripilus (Knight)

P. rubellicollis (Knight)

Tribe STENODEMINI

A Leptopterna dolabrata (Linnaeus)

L. ferrugata (Fallén)

Litomiris curtus (Knight)

L. debilis (Uhler)

Mimoceps insignis Uhler

A Megaloceroea recticornis (Geoffrey)

Stenodema pilosipes Kelton

S. trispinosa Reuter

S. vicina (Provancher)

Teratocoris caricis Kirkaldy

T. discolor Uhler

T. paludum Sahlberg

T. saundersi Douglas & Scott

Trigonotylus americanus Carvalho

T. coelestialium (Kirkaldy)

* T. longipes Slater & Wagner

T. viridis (Provancher)

Subfamily ORTHOTYLINAE

Tribe HALTICINI

Labops hesperius Uhler

L. hirtus Knight

L. tumidifrons Knight

L. verae Knight

Tribe ORTHOTYLINI

Aoplonema princeps (Uhler)

A. uhleri (Van Duzee)

A Blepharidopterus angulatus (Fallén)

B. provancheri (Burque)

B. ulmi (Knight)

Brooksetta inconspicua (Uhler)

B. incurva (Knight)

B. viridicata (Uhler)

Ceratocapsus apicatus Van Duzee

* C. cunealis Henry

C. drakei Knight

Cyrtorhinus caricis (Fallén)

Dichaetocoris piceicola (Knight)

Fieberocapsus flaveolus (Reuter)

Hadronema militare Uhler

H. simplex Knight

Ilnacora albifrons Knight

Ilnacorella argentata Knight

I. sulcata Knight

Labopidea lenensis (Lindberg)

L. nigrisetosa Knight

L. simplex (Uhler)

Lopidea dakota Knight

L. minor Knight

L. nigridea aculeata Van Duzee

L. nigridea nigridea Uhler

L. nigridea sericea Knight

* L. picta Knight

L. taurina Van Duzee

Macrotyloides vestitus (Uhler)

Mecomma angustatum (Uhler)

M. gilvipes (Stål)

Melanotrichus albocostatus (Van Duzee)

M. brindleyi Knight

M. coagulatus (Uhler)

M. eurotiae Knight

M. flavosparsus (Sahlberg)

M. leviculus Knight

M. mistus (Knight)

Noctuocoris fumidus (Van Duzee)

Orthotylus candidatus Van Duzee

O. contrastus Van Duzee

O. dorsalis (Provancher)

O. katmai (Knight)

O. neglectus Knight

O. pacificus Van Duzee

O. ute Knight

Paraproba nigrinervis Van Duzee

Parthenicus brindleyi Knight

P. brooksi Kelton

* P. pallidicollis Van Duzee

P. sabulosus Van Duzee

Pseudopsallus occidentalis Stonedahl & Schwartz

Sericophanes heidemanni Poppius

Slaterocoris atritibialis (Knight)

S. breviatus (Knight)

S. stygicus (Say)

Subfamily PHYLINAE

Tribe HALLODAPINI

Coquillettia insignis Uhler

O. montanus Knight

O. obliquus Uhler

Tribe LEUCOPHOROPTERINI

Tytthus pubescens (Knight)

Tribe PHYLINI

Atomoscelis modestus (Van Duzee)

Atractotomus cooperi Stonedahl

A Campylomma verbasci (Meyer-Dür)

Chaetophylidea moerens (Reuter)

Chlamydatus associatus (Uhler)

C. auratus Kelton

* C. brevicornis Knight

C. montanus Knight

C. obliquus (Uhler)

C. pallidicornis Knight

C. pulicarius (Fallén)

C. pullus (Reuter)

* C. schuhi Knight

Criocoris saliens (Reuter)

Dacota hesperia (Uhler)

Europiella artimisiae (Becker)

E. decolor (Uhler)

E. unipuncta Knight

Hoplomachus affiguratus (Uhler)

Knightomiroides ponderosae Stonedahl & Schwartz

A Lopus decolor (Fallén)

Macrotylus multipunctatus Van Duzee

Microphyllelus adustus binotatus Knight

Monosynamma bohemani (Fallén)

Phoenicocoris longirostris (Knight)

P. rostratus (Knight)

Phyllopidea montana Knight

P. picta Uhler

Plagiognathus alboradialis Knight

A P. chrysanthemi (Wolff)

P. flavidus Knight

P. fuscosus (Provancher)

P. fuscipes Knight

P. fuscotibialis Knight

P. medicagus Arrand

P. obscurus Uhler

* P. ribesi Kelton

P. shoshonea Knight

P. suffuscipennis Knight

* Pronotocrepis clavicornis Knight

Psallus aethiops (Zetterstedt)

P. falleni Reuter

P. parshleyi Knight

P. piceicola Knight

Sthenarus cuneotinctus (Van Duzee)

Tribe PILOPHORINI

Pilophorus americanus Poppius

P. diffusus Knight

* P. stonedahli Schuh & Schwartz

P. tibialis Van Duzee

P. vicarius Poppius

Family NABIDAE

Subfamily NABINAE

* Hoplistoscelis heidemanni (Reuter)

Nabicula americolimbata (Carayon)

N. nigrovittata nearctica Kerzhner

N. flavomarginata (Scholtz)

N. subcoleoptrata (Kirby)

N. vanduzeei (Kirkaldy)

Nabis roseipennis Reuter

N. rufusculus Reuter

N. alternatus alternatus Parshley

N. americoferus Carayon

N. inscriptus (Kirby)

Subfamily PROSTEMMATINAE

Pagasa fusca (Stein)

Superfamily TINGOIDEA

Family TINGIDAE

Acalypta cooleyi Drake

A. elegans Horvath

A. lillianis Torre Bueno

Corythaica venusta (Champion)

Corythuca distincta Osborn & Drake

C. elegans Drake

C. hewitti Drake

C. immaculata Osborn & Drake

C. juglandis (Fitch)

C. marmorata (Uhler)

C. mollicula Osborn & Drake

C. morrilli Osborn & Drake

C. padi Drake

C. pergandei Heidemann

C. salicata Gibson

Dictyla labeculata (Uhler)

* Gargaphia opacula (Uhler)

G. solani Heidemann

Hesperotingis antennata Parshley

H. occidentalis Drake

Melanorhopala clavata Stål

Physatocheila plexa (Say)

P. variegata Parshley

Infraorder LEPTOPODOMORPHA

Superfamily SALDOIDEA

Family SALDIDAE

Subfamily SALDINAE

Tribe SALDINI

Lampracanthia crassicornis (Uhler)

Salda anthracina Uhler

S. buenoi (McDunnough)

S. lugubris (Say)

S. obscura Provancher

S. provancheri Kelton & Lattin

Teloleuca bifasciata (Thomson)

Tribe SALDOIDINI

* Ioscytus politus (Uhler)

Micranthia bergrothi Jakovlev

M. fennica (Reuter)

M. humilis (Say)

M. quadrimaculata (Champion)

Saldula andrei Drake

S. basingeri Drake

S. comatula Parshley

S. dispersa (Uhler)

S. explanata (Uhler)

S. laticollis (Reuter)

S. opacula (Zetterstedt)

S. opiparia Drake & Hottes

S. pallipes (Fabricius)

S. palustris (Douglas)

S. saltatoria (Linnaeus)

Infraorder PENTATOMOMORPHA

Superfamily ARADOIDEA

Family ARADIDAE

Aneurus borealis Picchi

A. inconstans Uhler

A. simplex Uhler

Aradus abbas Bergroth

A. antennalis Parshley

A. approximatus Parshley

A. blaisdelli Van Duzee

A. coarctatus Heidemann

A. compressus Heidemann

A. debilis Uhler

A. depictus Van Duzee

A. falleni Stål

A. funestus Bergroth

A. heidemanni Bergroth

A. insolitus Van Duzee

A. intectus Parshley

A. kormilevi Heiss

A. lugubris Fallén

A. lugubris nigricornis Reuter

A. paganicus Parshley

A. parshleyi Van Duzee

A. parvicornis Parshley

A. persimilis Van Duzee

A. proboscideus Walker

A. quadrilineatus Say

A. similis Say

A. tuberculifer Kirby

A. vadosus Van Duzee

Superfamily COREOIDEA

Family ALYDIDAE

Alydus calcaratus (Linnaeus)

A. conspersus infuscatus Fracker

A. eurinus (Say)

A. scutellatus Van Duzee

Megalotomus quinquespinosus (Say)

Tollius curtulus (Stål)

Family COREIDAE

Anasa tristis (DeGeer)

Ceraleptus denticulatus Froeschner

C. pacificus Barber

Coriomeris humilis (Uhler)

Leptoglossus occidentalis Heidemann

Family RHOPALIDAE

A. scutatus (Stål)

Boisea rubrolineata (Barber)

Harmostes reflexulus (Say)

Liorhyssus hyalinus (Fabricius)

Stictopleurus punctiventris (Dallas)

S. viridicatus (Uhler)

Superfamily LYGAEOIDEA

Family BERYTIDAE

* Hoplinus echinata (Uhler)

Jalysus wickhami Van Duzee

Neoneides muticus (Say)

Family BLISSIDAE

Blissus occiduus Barber

Family CYMIDAE

Cymus coriacipennis (Stål)

C. luridus Stål

Family GEOCORIDAE

Geocoris atricolor Montandon

G. bullatus (Say)

G. discopterus Stål

G. howardi Montandon

G. limbatus Stål

G. pallens Stål

Family HETEROGASTRINAE

* Heterogaster behrensii (Uhler)

Family LYGAEIDAE

Subfamily ISCHNORHYNCHINAE

Kleidocerys franciscanus (Stål)

K. ovalis Barber

K. resedae (Panzer)

Subfamily LYGAEINAE

* Lygaeospilus brevipilus Scudder

Lygaeus kalmii kalmii Stål

Neacoryphus bicrucis (Say)

Subfamily ORSILLINAE

Nysius angustatus Uhler

N. fuscovittatus Barber

N. niger Baker

N. raphanus Howard

N. tenellus Barber

N. thymi (Wolff)

Neortholomus scolopax (Say)

Family OXYCARENIDAE

Crophius angustatus Van Duzee

C. bohemanni (Stål)

C. disconotus (Say)

C. ramosus Barber

Family PACHYGROWTHIDAE

Phlegyas annulicrus Stål

Family RHYPAROCHROMIDAE

Subfamily RHYPAROCHROMINAE

Tribe ANTILLOCORINI

* Botocudo modestus (Barber)

Tribe DRYMINI

Eremocoris borealis (Dallas)

E. obscurus Van Duzee

** Gastrodes intermedius Usinger

G. pacificus (Provancher)

Scolopostethus atlanticus Horvath

S. pacificus Barber

S. thomsoni Reuter

Tribe GONIANOTINI

Emblethis vicarius Horvath

M. sodalicius (Uhler)

Trapezonotus arenarius (Linnaeus)

Tribe MEGALONOTINI

A Megalonotus sabulicola (Thomson)

Sphragisticus nebulosus (Fallén)

Tribe MYODOCHINI

Kolenetrus plenus (Distant)

Ligyrocoris diffusus (Uhler)

L. latimarginatus Barber

L. sylvestris (Linnaeus)

Pseudocnemodus canadensis (Provancher)

* Sisamnes claviger (Uhler)

Slaterobius insignis (Uhler)

Zeridoneus costalis (Van Duzee)

Tribe OZOPHORINI

* Ozophora occidentalis Slater

Tribe RHYPAROCHROMINI

* Cordillonotus stellatus Scudder

Peritrechus convivus (Stål)

P. fraternus Uhler

P. tristis Van Duzee

Tribe STYGNOCORINI

A Stygnocoris rusticus (Fallén)

A S. sabulosus (Schilling)

Tribe UDEOCORINI

* Neosuris castanea (Barber)

Superfamily PENTATOMOIDEA

Family ACANTHOSOMATIDAE

Elasmostethus cruciatus (Say)

Elasmucha lateralis (Say)

Family CYDNIDAE

Amnestus pallidus Zimmer

Sehirus cinctus albonotatus Dallas

Family PENTATOMIDAE

Subfamily ASOPINAE

Perillus bioculatus (Fabricius)

P. exaptus (Say)

Podisus brevispinus Phillips

P. placidus Uhler

P. serieventris Uhler

Rhacognathus americanus Stål

Zicrona caerulea (Linnaeus)

Subfamily PENTATOMIDAE

Tribe HYALINI

Brochymena affinis Van Duzee

B. quadripustulata (Fabricius)

Tribe PENTATOMINI

Acrosternum hilare (Say)

Aelia americana Dallas

Banasa dimidiata (Say)

B. sordida (Uhler)

Chlorochroa congrua Uhler

C. granulosa (Uhler)

C. ligata (Say)

C. opuntiae Esselbaugh

C. rossiana Buxton & Thomas

C. uhleri (Stål)

Codophila remota (Horvath)

Coenus delius (Say)

Cosmopepla bimaculata (Thomas)

C. intergressus (Uhler)

Dendrocoris pini Montandon

Euschistus conspersus Uhler

E. servus euschistoides (Vollenhoven)

E. tristigmus luridus Dallas

E. variolarius (Palisot)

Holcostethus abbreviatus Uhler

H. limbolarius (Stål)

H. piceus (Dallas)

H. tristis (Van Duzee)

Neottiglossa trilineata (Kirby)

N. tumidifrons Downes

N. undata (Say)

Prionosoma podopoides Uhler

Tepa rugulosa (Say)

Thyanta pallidovirens (Stål)

Trichopepla atricornis Stål

T. aurora Van Duzee

T. dubia (Dallas)

Tribe SCIOCORINI

Sciocoris microphthalmus Flor

Family SCUTELLERIDAE

Subfamily EURYGASTRINAE

Eurygaster amerinda Bliven

Phimodera binotata (Say)

Vanduzeeina balli (Van Duzee)

V. borealis Van Duzee

Subfamily PACHYCORINAE

Homaemus aeneifrons consors Uhler

H. bijugis (Uhler)

* H. parvulus (Germar)

Family THYREOCORIDAE

Corimelaena extensa Uhler

C. nigra Dallas

C. pulicaria (Germar)

Galgupha nitiduloides nitiduloides (Wolff)

G. ovalis Hussey

Superfamily PIESMATOIDEA

Family PIESMATIDAE

Piesma cinereum (Say)

P. explanatum McAtee